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What causes the following warning: "WARNING: PDB2PQR was unable to assign charges to the following atoms"?

Some of the protonation states derived from the [PROPKA]] results are not supported in the requested forcefield and thus PDB2PQR is unable to get charges and radii for that state. PDB2PQR currently supports the following states as derived from PROPKA:
Protonation StateAMBER SupportCHARMM SupportPARSE Support
Neutral N-TerminusNoNoYes
Neutral C-TerminusNoNoYes
Neutral ARGNoNoNo
Neutral ASPYes†YesYes
Negative CYSYes†NoYes
Neutral GLUYes†YesYes
Neutral HISYesYesYes
Neutral LYSYes†NoYes
Negative TYRNoNoYes
† - only if residue is not a terminal residue; if the residue is terminal it will not be set to this state